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Contributors MAM-C, JRG-L and MFG-E contributed to the conception and design of the research, analysis and interpretation of the data, draft and revision of the manuscript. LO-F and MC-J performed the experiments and analysed the data. The rest of the authors: NO-C, MJC-P, GE, GG-G, MAG-G, ACB-M, RS, PF, MRC, TC, SC, JS-B and JM contributed with samples and patient information. All authors read and approved the final version.
Funding This work was supported by Fondo de Investigaciones Sanitarias (10/1701), Fondos FEDER, Plan Andaluz de Investigación (CTS-0197 and CTS-180), Red Enfermedades Inflamatorias y Reumáticas and Consejería de Salud de la Junta de Andalucía (0260/08). LOF is the recipient of a fellowship (FI11/00547).
Competing interests None.
Patient consent Obtained.
Ethics approval The local ethical committees of all the institutions involved.
Provenance and peer review Not commissioned; externally peer reviewed.
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